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  2. Staff Directory
  3. Principal Investigators Directory
  4. Research Areas
  5. The Tumor Microenvironment

Investigator Directory

The Tumor Microenvironment

The molecular makeup and biological properties of cancer cells can vary significantly, even within the same tumor. Tumors are highly heterogeneous and a few specialized cells drive the growth of the entire tumor. Ongoing studies seek to understand the molecular differences between the various types of cells in a tumor. We invite you to meet some of our principal investigators conducting research in this area.


  • Clint Allen
    Acting Chief
    Clint T. Allen, M.D.
    clint.allen@nih.gov
  • Stefan  Ambs, Ph.D., M.P.H.
    Senior Investigator
    Stefan Ambs, Ph.D., M.P.H.
    ambss@mail.nih.gov
  • Takeo Fujii
    Physician-Scientist Early Investigator
    Takeo Fujii, M.D., M.P.H.
    takeo.fujii@nih.gov
  • Rosandra N. Kaplan, M.D.
    Senior Investigator
    Rosandra N. Kaplan, M.D.
    kaplanrn@mail.nih.gov
  • Troy McEachron PhD
    Investigator
    Troy A. McEachron, Ph.D.
    troy.mceachron@nih.gov
  • Meera Murgai, Ph.D.
    Stadtman Investigator
    Meera Murgai, Ph.D.
    meera.murgai@nih.gov
  • David D. Roberts, Ph.D.
    Senior Investigator
    David D. Roberts, Ph.D.
    droberts@mail.nih.gov
  • William G. Stetler-Stevenson, M.D., Ph.D.
    Senior Investigator
    William G. Stetler-Stevenson, M.D., Ph.D.
    sstevenw@mail.nih.gov
  • Anish  Thomas, MBBS, M.D.
    Senior Investigator
    Anish Thomas, MBBS, M.D.
    anish.thomas@nih.gov
  • Philip  Tofilon, Ph.D.
    Senior Investigator
    Philip Tofilon, Ph.D.
    tofilonp@mail.nih.gov
  • Giovanna Tosato, M.D.
    Senior Investigator
    Giovanna Tosato, M.D.
    tosatog@mail.nih.gov
  • Li  Yang, Ph.D.
    Senior Investigator
    Li Yang, Ph.D.
    yangl3@mail.nih.gov
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